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söndag 27 maj 2018

APOBEC3C (Kr.22q13.1), A3C, PBI, ARP5, ARDC2, ARDC4

APOBEC3C,  A3C,  ARP5

Samassa Kr.22 klusterissa seitsemän samantapaisen geenin kanssa geeniduplikaatiolla muodostuneen geenin tai pseudogeenin kanssa, ne koodaavat rakenteellisesti ja toiminnallisesti samantapaisia sydidiini C- deaminaaseja kuin APOBEC1 ja tuottavat urasiilia U. On ajateltu näiden proteiinien muovaaman entsymaattisesti RNA:ta ja osallistuvan kasvuun tai solusyklin kontrolliin.Tätä geeniä esiintyy yleisesti lymfasolmukkeissa, testiksessä ja 24 muusas kudoksessa.Tämän geenin deleetio ei näytä olevan hyödyksi ainakaan rintasyövässä. retrovirusvastaista merkitsytä lienee tälläkin Apobec3C- lajilla, vaikka   ei niin selvästi kuin jollain muulla tässä klusterissa. Sisältää kuitenkin  merkitseviä motiiveja kuten f-y-v-e-

Mitään silmiinpistävän suurta  muuta erityismerkitystä  en löydä tästä geenist, mikä ei olisi muiten APOBEC3- proteiinien kattamatonta. Molekyyli on  190 aminohappoa sisältävä.
  • https://www.ncbi.nlm.nih.gov/gene/27350 Also known as A3C; PBI; ARP5; ARDC2; ARDC4; APOBEC1L; bK150C2.3 Summary This gene is a member of the cytidine deaminase gene family. It is one of seven related genes or pseudogenes found in a cluster thought to result from gene duplication, on chromosome 22. Members of the cluster encode proteins that are structurally and functionally related to the C to U RNA-editing cytidine deaminase APOBEC1. It is thought that the proteins may be RNA editing enzymes and have roles in growth or cell cycle control. [provided by RefSeq, Jul 2008] Expression Ubiquitous expression in lymph node (RPKM 33.1), testis (RPKM 26.8) and 24 other tissues See more Orthologs all

Preferred Names

DNA dC->dU-editing enzyme APOBEC-3C
Names
apolipoprotein B editing enzyme catalytic polypeptide-like 3C
apolipoprotein B mRNA editing enzyme, catalytic polypeptide-like 3C
phorbolin I
probable DNA dC->dU-editing enzyme APOBEC-3C

Conserved Domains (3) summary

cd01283
Location:8 → 158
cytidine_deaminase; Cytidine deaminase zinc-binding domain. These enzymes are Zn dependent. The zinc ion in the active site plays a central role in the proposed catalytic mechanism, activating a water molecule to form a hydroxide ion that performs a nucleophilic attack on ...
pfam05240
Location:129 → 174
APOBEC_C; APOBEC-like C-terminal domain
pfam08210
Location:17 → 187
APOBEC_N; APOBEC-like N-terminal domain
      • PEPTIDE sequence and history

https://www.ncbi.nlm.nih.gov/protein/NP_055323.2
  • DNA C-to U -editing enzyme APOBEC-3C [Homo sapiens]
    NCBI Reference Sequence: NP_055323.2
  • LOCUS       NP_055323                190 aa            linear   PRI 01-APR-2018
    DEFINITION  DNA dC->dU-editing enzyme APOBEC-3C [Homo sapiens].
    ACCESSION   NP_055323
    VERSION     NP_055323.2
    DBSOURCE    REFSEQ: accession NM_014508.2
    KEYWORDS    RefSeq.
    SOURCE      Homo sapiens (human)
      ORGANISM  Homo sapiens
                Eukaryota; Metazoa; Chordata; Craniata; Vertebrata; Euteleostomi;
                Mammalia; Eutheria; Euarchontoglires; Primates; Haplorrhini;
                Catarrhini; Hominidae; Homo.
    REFERENCE   1  (residues 1 to 190)
      AUTHORS   Chen Z, Eggerman TL, Bocharov AV, Baranova IN, Vishnyakova TG,
                Kurlander R and Patterson AP.
      TITLE     Heat shock proteins stimulate APOBEC-3-mediated cytidine
                deamination in the hepatitis B virus
      JOURNAL   J. Biol. Chem. 292 (32), 13459-13479 (2017)
       PUBMED   28637869
      REMARK    GeneRIF: Data suggest that heat shock proteins, in particular
                Hsp90, stimulate APOBEC3-mediated DNA deamination activity toward
                hepatitis B viral DNA, suggesting a potential physiological role in
                mutagenesis/carcinogenesis and viral innate immunity; Hsp90
                stimulates deamination activity of APOBEC3G, APOBEC3B, and APOBEC3C
                during co-expression in human liver HepG2 cells.
    REFERENCE   2  (residues 1 to 190)
      AUTHORS   Jaguva Vasudevan AA, Hofmann H, Willbold D, Haussinger D, Koenig BW
                and Munk C.
      TITLE     Enhancing the Catalytic Deamination Activity of APOBEC3C Is
                Insufficient to Inhibit Vif-Deficient HIV-1
      JOURNAL   J. Mol. Biol. 429 (8), 1171-1191 (2017)
       PUBMED   28315663
      REMARK    GeneRIF: Antiviral functions of APOBEC3C against HIV-1 and APOBEC3C
                binding capacity
    REFERENCE   3  (residues 1 to 190)
      AUTHORS   Middlebrooks CD, Banday AR, Matsuda K, Udquim KI, Onabajo OO,
                Paquin A, Figueroa JD, Zhu B, Koutros S, Kubo M, Shuin T, Freedman
                ND, Kogevinas M, Malats N, Chanock SJ, Garcia-Closas M, Silverman
                DT, Rothman N and Prokunina-Olsson L.
      TITLE     Association of germline variants in the APOBEC3 region with cancer
                risk and enrichment with APOBEC-signature mutations in tumors
      JOURNAL   Nat. Genet. 48 (11), 1330-1338 (2016)
       PUBMED   27643540
    REFERENCE   4  (residues 1 to 190)
      AUTHORS   Wittkopp CJ, Adolph MB, Wu LI, Chelico L and Emerman M.
      TITLE     A Single Nucleotide Polymorphism in Human APOBEC3C Enhances
                Restriction of Lentiviruses
      JOURNAL   PLoS Pathog. 12 (10), e1005865 (2016)
       PUBMED   27732658
      REMARK    GeneRIF: our results suggest that APOBEC3C is in fact involved in
                protecting hosts from lentiviruses.
                Publication Status: Online-Only
    REFERENCE   5  (residues 1 to 190)
      AUTHORS   Zhang Y, Delahanty R, Guo X, Zheng W and Long J.
      TITLE     Integrative genomic analysis reveals functional diversification of
                APOBEC gene family in breast cancer
      JOURNAL   Hum. Genomics 9, 34 (2015)
       PUBMED   26682542
      REMARK    GeneRIF: These results suggest that functional potential of
                APOBEC3B and APOBEC3C involved in cancer mutagenesis is associated
                with estrogen receptor status.
                Publication Status: Online-Only
    REFERENCE   6  (residues 1 to 190)
      AUTHORS   Mariani R, Chen D, Schrofelbauer B, Navarro F, Konig R, Bollman B,
                Munk C, Nymark-McMahon H and Landau NR.
      TITLE     Species-specific exclusion of APOBEC3G from HIV-1 virions by Vif
      JOURNAL   Cell 114 (1), 21-31 (2003)
       PUBMED   12859895
    REFERENCE   7  (residues 1 to 190)
      AUTHORS   Wedekind JE, Dance GS, Sowden MP and Smith HC.
      TITLE     Messenger RNA editing in mammals: new members of the APOBEC family
                seeking roles in the family business
      JOURNAL   Trends Genet. 19 (4), 207-216 (2003)
       PUBMED   12683974
      REMARK    Review article
                Erratum:[Trends Genet. 2003 Jul;19(7):369]
    REFERENCE   8  (residues 1 to 190)
      AUTHORS   Jarmuz A, Chester A, Bayliss J, Gisbourne J, Dunham I, Scott J and
                Navaratnam N.
      TITLE     An anthropoid-specific locus of orphan C to U RNA-editing enzymes
                on chromosome 22
      JOURNAL   Genomics 79 (3), 285-296 (2002)
       PUBMED   11863358
    REFERENCE   9  (residues 1 to 190)
      AUTHORS   Dunham I, Shimizu N, Roe BA, Chissoe S, Hunt AR, Collins JE,
                Bruskiewich R, Beare DM, Clamp M, Smink LJ, Ainscough R, Almeida
                JP, Babbage A, Bagguley C, Bailey J, Barlow K, Bates KN, Beasley O,
                Bird CP, Blakey S, Bridgeman AM, Buck D, Burgess J, Burrill WD,
                O'Brien KP et al.
      TITLE     The DNA sequence of human chromosome 22
      JOURNAL   Nature 402 (6761), 489-495 (1999)
       PUBMED   10591208
      REMARK    Erratum:[Nature 2000 Apr 20;404(6780):904]
    REFERENCE   10 (residues 1 to 190)
      AUTHORS   Madsen P, Anant S, Rasmussen HH, Gromov P, Vorum H, Dumanski JP,
                Tommerup N, Collins JE, Wright CL, Dunham I, MacGinnitie AJ,
                Davidson NO and Celis JE.
      TITLE     Psoriasis upregulated phorbolin-1 shares structural but not
                functional similarity to the mRNA-editing protein apobec-1
      JOURNAL   J. Invest. Dermatol. 113 (2), 162-169 (1999)
       PUBMED   10469298
    COMMENT     REVIEWED REFSEQ: This record has been curated by NCBI staff. The
                reference sequence was derived from BC011739.2.
                On Sep 16, 2002 this sequence version replaced NP_055323.1.
                
                Summary: This gene is a member of the cytidine deaminase gene
                family. It is one of seven related genes or pseudogenes found in a
                cluster thought to result from gene duplication, on chromosome 22.
                Members of the cluster encode proteins that are structurally and
                functionally related to the C to U RNA-editing cytidine deaminase
                APOBEC1. It is thought that the proteins may be RNA editing enzymes
                and have roles in growth or cell cycle control. [provided by
                RefSeq, Jul 2008].
                
                Publication Note:  This RefSeq record includes a subset of the
                publications that are available for this gene. Please see the Gene
                record to access additional publications.
                
                ##Evidence-Data-START##
                Transcript exon combination :: BC011739.2, SRR1163655.295268.1
                                               [ECO:0000332]
                RNAseq introns              :: single sample supports all introns
                                               SAMEA1965299, SAMEA1966682
                                               [ECO:0000348]
                ##Evidence-Data-END##
    FEATURES             Location/Qualifiers
         source          1..190
                         /organism="Homo sapiens"
                         /db_xref="taxon:9606"
                         /chromosome="22"
                         /map="22q13.1"
         Protein         1..190
                         /product="DNA dC->dU-editing enzyme APOBEC-3C"
                         /note="probable DNA dC->dU-editing enzyme APOBEC-3C;
                         phorbolin I; apolipoprotein B editing enzyme catalytic
                         polypeptide-like 3C; apolipoprotein B mRNA editing enzyme,
                         catalytic polypeptide-like 3C"
                         /calculated_mol_wt=22695
         Region          8..158
                         /region_name="cytidine_deaminase"
                         /note="Cytidine deaminase zinc-binding domain. These
                         enzymes are Zn dependent. The zinc ion in the active site
                         plays a central role in the proposed catalytic mechanism,
                         activating a water molecule to form a hydroxide ion that
                         performs a nucleophilic attack on...; cd01283"
                         /db_xref="CDD:238610"
         Region          17..187
                         /region_name="APOBEC_N"
                         /note="APOBEC-like N-terminal domain; pfam08210"
                         /db_xref="CDD:285428"
         Site            order(32,34,55,57,66..68,97,100)
                         /site_type="active"
                         /db_xref="CDD:238610"
         Site            order(66..68,96..97,100)
                         /site_type="active"
                         /note="catalytic motif [active]"
                         /db_xref="CDD:238610"
         Site            order(66,68,97,100)
                         /site_type="other"
                         /note="Zn binding site [ion binding]"
                         /db_xref="CDD:238610"
         Region          129..174
                         /region_name="APOBEC_C"
                         /note="APOBEC-like C-terminal domain; pfam05240"
                         /db_xref="CDD:283020"
         CDS             1..190
                         /gene="APOBEC3C"
                         /gene_synonym="A3C; APOBEC1L; ARDC2; ARDC4; ARP5;
                         bK150C2.3; PBI"
                         /coded_by="NM_014508.2:104..676"
                         /db_xref="CCDS:CCDS13983.1"
                         /db_xref="GeneID:27350"
                         /db_xref="HGNC:HGNC:17353"
                         /db_xref="MIM:607750"
    ORIGIN      
            1 mnpqirnpmk amypgtfyfq fknlweandr netwlcftve gikrrsvvsw ktgvfrnqvd
           61 sethchaerc flswfcddil spntkyqvtw ytswspcpdc agevaeflar hsnvnltift
          121 arlyyfqypc yqeglrslsq egvaveimdy edfkycwenf vyndnepfkp wkglktnfrl
          181 lkrrlreslq
    //

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